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BFG I cant speak for aslavatortin . but I think you want it before the skimmer since the skimmer would be removing the food that the sponges would need. thats my .02 c. if you look at his profile he hasnt logged on since 2009. but i could be wrong. one of the things that steve ( i am not trying to speak for steve ) wrote in his books was that the sponges might foul with large particulate matter and be best in low flow areas. thats what I remember from reading his books many years ago. I think the 404 has a large flow so a conundrum. Many types of sponges so many possibilities. you will probably have to experiment . I do believe in the different zones to be necessary yet may not be practical to have in the small footprints that we have in our homes. One of the things that comes to mind is Koi filtration. to have clear water you need large settling pond to pump up to a planted section to absorb nutrients which end up needing a large surface area to filter the pond.
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